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Using cards in Story Mode and Event Battles give those cards experience with each win. Starting in Version 3.2 that experience also counts toward each player's Level.
Once a card is Mastered, you gain Mastery Points for each of the card's colors based on the card's rarity. Colorless cards give 1/2 of the Mastery Points for that rarity, but for all 5 colors. You tier up in each color separately.
Once a card is mastered you '''CANNOT''' gain any more experience or Mastery Points from that card.
